The Networks International Council provides for exchange of information and collaborative activities with some other networks internationally. Members are directors of their Networks, and meet twice yearly by video-conference .

Current membership is:

	Quantifying and Understanding the Earth System (QUEST, UK): Director Colin Prentice
	National Evolutionary Synthesis Center (NESCent, USA): Director Kathleen Smith
	National Center for Ecological Analysis and Synthesis (NCEAS, USA): Director Jim Reichman
	ARC-NZ Research Network for Vegetation Function: Convenor Mark Westoby